Loeffler succeeded in cultivating C. diphtheriae, the diphtheria bacillus. He reproduced the characteristic membrane by swabbing the mucous membranes of various animals with pure cultures of the bacillus. In this pape…
J. S. Anderson, F. C. Happold, J. W. McLeod, and J. G. Thomson were the first to distinguish the gravis, mitis, and intermediate types of C. diphtheriae.